Notes on Afrotropical and Palearctic species of the genus Thinophilus

Wahlberg (Diptera: Dolichopodidae) with descriptions of new species

Igor Ya. GRICHANOV

Grichanov, I.Ya. Notes on Afrotropical and Palearctic species of

the genus Thinophilus Wahlberg (Diptera: Dolichopodidae) with

descriptions of new species

T. munroi setiscutellatus ssp.n. from Namibia, T. ciliventris

sp.n. from South Africa, Botswana, Angola, and Nigeria are described.

New records for known Afrotropical and Palearctic species are given.

Catalogue and key to species of Thinophilus from both Regions are also

present.

I.Ya. Grichanov, All-Russian Institute of Plant Protection,

Podbelskogo 3, St.Petersburg-Pushkin, 189620, Russia.

Key words: Diptera, Dolichopodidae, Thinophilus, key,

Tropical Africa, Palearctic Region

Introduction

Widely distributed halophilous species of the genus Thinophilus

Wahlberg are confined mostly to arid zones and also to sea coast in

many Regions of the world. Concerning the fauna of the Old World, I

regard that one of the centers of the genus diversity is located at

the border dividing Afrotropical and Palearctic Regions. Totally 40

species are known today from the two Regions. 13 species are recorded

only from Afrotropical Region, 10 species - only from Palearctic

Region, whereas 16 Afrotropical and Palearctic species are located in

Saharan-Arabian Transitional Zone. I conditionally refer to this Zone

the following countries mentioned in the Catalogs (Dyte & Smith,

1980; Negrobov, 1991): Morocco, Algeria, Tunisia, Egypt, Sudan,

Ethiopia, Djibouti, Somalia, and Yemen.

The most interest distribution of species are as follows: T.

indigenus Becker - Mongolia, Iran, Turkey, Algeria, Egypt, Cape Verde

Is., Ethiopia, South Yemen, Nigeria (!), Benin (!), Ghana (!), Zaire,

Tanzania (!), Madagascar, Angola (!), Namibia (!), South Africa (!),

Swaziland (!); Oriental Region; T. mirandus Becker - Algeria, Morocco,

Tanzania (!); T. palpatus Parent - Ethiopia, Nigeria, Angola (!),

Namibia (!), Botswana (!). Several Palearctic species had doubtful

records from Tropical Africa, and some Saharan species were also

recorded from South Europe and/or Central Asia. I would expect that

many species had wider distribution than area known up-to-day. These

data induced me to compile the joint key to all Afrotropical and

Palearctic species of Thinophilus (see below).

All the species described and recorded are referred to the

subgenus Thinophilus s.s. T. prudens Curran, probably, cannot be

united with T. versutus Walker in the subgenus Schoenophilus Mik in

spite of the presence of 4 dorsocentral setae on mesonotum in both

species (compare descriptions in Curran, 1926 and Meuffels &

Grootaert, 1984). On the other hand, T. munroi munroi Curran and T.

munroi setiscutellatus ssp.n. should be possibly separated from other

species in new subgenus. Some of the species are known from females,

others - only from males. Therefore, several species cannot be surely

identified without opposite sex of the same series.

Treating unidentified material from the collections of the

Natural History Museum, London [NHML], the Hungarian Natural History

Museum [HNHM], and Lund University, Sweden [Lund], I found many

interest species. In this paper descriptions of T. munroi setiscutellatus

ssp.n. from Namibia, T. ciliventris sp.n. from South Africa, Botswana,

Angola, and Nigeria, and new records for known African and Palearctic

species are given. Catalogue and key to known species of Thinophilus

from both Regions are also present.

Holotypes and paratypes of the new species are deposited in the

Natural History Museum (London). Some specimens collected by author

are conserved in the collection of Voronezh University, Russia [VU].

List of Afrotropical species of Thinophilus unknown from

Palearctic and Transitional Afro-Palearctic Regions

(for references see Dyte & Smith, 1980)

annulitarsis Parent, 1936:323 - Tanzania

aquaticus Becker, 1914:125 - Kenya, Madagascar

bipunctatus Curran, 1926:27 - South Africa, Zaire

calopus Loew, 1852:659 - Mozambique, Zaire

capensis Curran, 1926:26 - South Africa, Zaire

ciliventris sp.n. - South Africa, Botswana, Angola, Nigeria.

imperialis Curran, 1924:228 (Nematoproctus) - South Africa,

Botswana (!), Congo, Ghana, Nigeria, Zaire

munroi munroi Curran, 1926:20 - South Africa

munroi setiscutellatus ssp.n. - Namibia

prudens Curran, 1926:30 - South Africa, Angola (!), Ghana, Zaire,

Senegal (!)

quadrisetus Parent, 1936:324 - Tanzania, Zaire

rex Curran, 1926:23 - South Africa, Zaire

splendidus Vanschuytbroeck, 1951:80 - Zaire

virgatus Curran, 1926:22 - South Africa

List of Palearctic species of Thinophilus unknown from

Afrotropical and Transitional Afro-Palearctic Regions

(for references see Negrobov, 1991)

bicalcaratus Negrobov, 1971:901 - Uzbekistan, Tadjikistan, Turkmenia

brevicilius Negrobov, 1971:901 - Uzbekistan, Tadjikistan, Kirgizia (!)

longipilus Negrobov, 1971:902 - Russian Far East

neptunus Frey, 1915:78 - Sweden

ornatus Negrobov et Grichanov, 1982:105 - Tadjikistan

pollinosus Loew, 1870:58 - China, Mongolia, Tadjikistan

ruficornis Haliday, 1838:184 (Medeterus) - whole Europe, West Siberia

(Omsk Region, Novosibirsk Region), North Kazakhstan, Kirgizia,

Mongolia, North China

= maculicornis Zetterstedt, 1843:474 (Rhaphium)

setosus Negrobov, 1979:433 - Mongolia

spinitarsis Becker, 1907:315 - China, Iran, Tadjikistan (!), South

Ukraine (Kherson Region)

vanschuytbroecki Negrobov, 1971:902 - Azerbaidjan, Turkmenistan,

Afghanistan

List of Palearctic and Afrotropical species of Thinophilus

found in Transitional Afro-Palearctic Region

(for references see Dyte & Smith, 1980; Negrobov, 1991)

achylleus Mik, 1900:73 - Egypt, Tunisia, Italy, Spain

= albidus Macquart, 1849 (Hydrophorus)

argyropalpis Becker, 1910:25 - Egypt, Tunisia, Algeria, South Arabia,

Iran, Uzbekistan, Turkmenia, Tadjikistan (!), Kazakhstan, South

Russia (Volgograd Region), Ukraine (Odessa Region)

atritarsis Parent, 1929:53 - Sudan

flavipalpis Zetterstedt, 1843:472 (Rhaphium) - Egypt, West Europe

(except North), Ukraine (Odessa Region, Zaporozh'e Region,

Crimea), South Russia (Rostov Region), Caucasus, Kazakhstan,

Kirgizia, Mongolia; ?Mozambique

indigenus Becker, 1902:481 - Mongolia, Iran, Turkey, Algeria,

Egypt, Cape Verde Is., Ethiopia, South Yemen, Nigeria (!), Benin

(!), Ghana (!), Zaire, Tanzania (!), Madagascar, Angola (!),

Namibia (!), South Africa (!), Swaziland (!); India, Nepal,

Malaya, Borneo, Philippines, Taiwan

maculatus Parent, 1929:50 - Sudan

mirandus Becker, 1907:112 - Algeria, Morocco, Tanzania (!)

modestus Becker, 1902:50, 75 - Egypt, ?Austria

ochripalpis Becker, 1910:139 - South Yemen, Somalia

palpatus Parent, 1930:101 - Ethiopia, Nigeria, Angola (!), Namibia

(!), Botswana (!)

promotus Becker, 1910:23 - South Yemen, Djibouti

quadrimaculatus Becker, 1902:49 - Egypt, Tunisia, Algeria,

Tadjikistan, Iran; ?Zaire

setulipalpis Bezzi, 1906:302 - Ethiopia (Eritrea), Zaire, ?Italy

spinulosus Parent, 1929:48 - Sudan, Somalia, Nigeria

tinctus Parent, 1929:51 - Sudan, ?Madagascar, Zaire

(Schoenophilus) versutus Haliday in Walker, 1851:192 - Algeria,

Morocco, Central and South Europe, Ukraine (Lugansk Region)

= maculipennis Strobl, 1899:123 (Pseudacropsilus)
 
 
 
 

Descriptions and new records

Thinophilus quadrimaculatus Becker

Material examined. Male, South Tadjikistan: Tigrovaya Balka,

tugai, 21.VII.1984, Grichanov [Author's Coll.].

Diagnosis. The species is well recognizable by four black lateral

spots on mesonotum and smoky spot near the end of R2+3 and R4+5 on

wing.

Distribution. Egypt, Tunisia, Algeria, ?Zaire; Tadjikistan.

Thinophilus indigenus Becker

Material examined. 2 males, 1 female, Swaziland: 3 km N Simunye,

26o11' S, 31o57' E, 27.X.1994, loc. 36, leg. R. Danielsson [Lund].

Female, RSA: Cape Prov., Cedarberg, 3 km ESE Kriedowkrans, 32o22' S,

18o59' E, 350 m, 06.X.1994, loc. 10, leg. R. Danielsson [Lund].

Female, Gambia: Bakau, Cape St. Mary at Sun Wing Hotel, swept in veg.

along the beach, 5.XI.1977, UTM 28PCK1991, Loc. 26 / Lund Univ., Syst.

Dept., Sweden Gambia/Senegal. Nov. 1977, Cederholm - Danielsson -

Hammarstedt - Hedquist - Samuelsson. Male & female, Manuara Steppe,

1979.IV.20, leg. Eory-Sipos [HNHM]. 5 males & 3 females, S.W. Africa

(W1): Nr. Onseepkans, Orange R. banks, 8-10.I.1972 / Southern African

Exp. B.M. 1972-1. 2 males, Angola (A11): Bruco, 26.II-2.III.1972 /

Southern African Exp. B.M. 1972-1. 2 females, Angola (A32):

Cachoeiras, 20 mls. SW Gabela, 18-19.III.1972 / Southern African Exp.

B.M. 1972-1. Female, S.W. Africa (W52): Swakop R., 3 mls. S.

Okahandja, 7.IV.1972 / Southern African Exp. B.M. 1972-1. Female, S.

Africa (S24): C.P., Dwyka R., Merwevilk-Koup rd., 2.I.1972 / Southern

African Exp. B.M. 1972-1. Female, Nigeria: Ile-Ife, W State, 15 Aug.

1971, Col. J.T. Medler / DD-46 [NHML]. Female, Tanganyika T.:

Morogoro, 29.I.1917, A. Loveridge / Pres. by Imp. Inst. Ent. Brit.

Mus. 1932-301. Female, Gold Coast: Taimong nr. Accra, 18.III.1922, Dr.

J.W. Scott Macfie / From edge of washing pool / Pres. by Imp. Bur.

Ent. Brit. Mus. 1923-6.

Diagnosis. The species can be recognized by five dark spots on

mesonotum and simple tarsi.

Distribution. Mongolia, Iran, Turkey, Algeria, Egypt, Cape Verde

Is., Ethiopia, South Yemen, Nigeria (!), Benin (!), Ghana (!), Zaire,

Tanzania (!), Madagascar, Angola (!), Namibia (!), South Africa (!),

Swaziland (!); India, Nepal, Malaya, Borneo, Philippines, Taiwan.

Thinophilus rex Curran

Material examined. 2 females, S. Africa: R.E. Turner, Brit. Mus.

1932-145 / Cape Province, Heidelberg dist., Breede River, 6.II.1932.

Diagnosis. T. rex differs from other species with maculated

thorax by hyaline wing, four spots on mesonotum and two spots on

scutellum.

Distribution. South Africa, Zaire.

Thinophilus prudens Curran

(Fig. 5)

Material examined. Male & female, Senegal: in forest 1.5 km NE

Djibelor ca. 6.5 km SW Ziguinchor, 8.III.1977, At light 19.00-21.30,

Loc. No. 23A, UTM 28PCJ575885. / Lund Univ., Syst. Dept., Sweden

Gambia/Senegal. Febr.-March 1977, Cederholm - Danielsson - Larsson -

Mirestrom - Norling - Samuelsson. Male. Angola: Huila District, J.

Balfour-Browne, B.M. 1954-797 / Rio Coroca, 8 mls. N. of Porto

Alexandre, 23. VI.1954, s.l. flying round pools / Stn. No. 290.

Female, Angola (A2): Rocadas, R. Cumene, 19-22.II.1972 / at light /

Southern African Exp. B.M. 1972-1;

Description (male from Angola). Frons bronze-green. Black

postocular setae in upper row on either side present, one black

vertical seta laterally on frons, one pair of ocellar setae, one pair

of occipital setae. Ventral postcranium covered with dirty-white

irregular postocular setae. Face metallic, grey pollinose. Clypeus

nearly twice as wide as high. Antenna short, mostly dark-brown, scape

and pedicel ventrally yellow, first flagellomere rounded, with short

terminal hairs. Arista dorsal, bare, thick at base. Length ratio of

first flagellomere to arista, 7 : 36. Palpus yellow, nearly twice as

high as clypeus, greyish pollinose, with sparse dark hairs mainly

along internal margin. Proboscis black, with white hairs.

Thorax bronze-green, grey pollinose, without dark spots. Pleura

densely whitish pollinose. Four strong dorsocentral bristles with

short hair in front of the first one somewhat laterally; no

acrostichals. Propleura with a few white hairs. Scutellum with 2

strong black setae.

Legs yellow, tarsi brown except base of basitarsomeres. Fore coxa

yellow, with short white hairs. Middle and hind coxae dark except

apex, grey pollinose, with a few short light hairs. Fore femora with

ventral row of very fine hairs, twice as long as diameter of femora.

Fore tibia with fine postero-dorsal seta. Length ratio of fore coxa to

femora to tibia to tarsus (segments from first to fifth) - 30 : 45 :

45 : 17 : 6 : 6 : 6 : 7. Middle femora with a row of ventral hairs in

basal half, as long as diameter of femora. Middle tibia with one

antero-dorsal, one postero-dorsal and row of fine ventral hairs in

apical 2/3, as long as tibia diameter. Length ratio of middle coxa to

femora to tibia to tarsus (segments from first to fifth) - 25 : 60 :

60 : 21 : 12 : 9 : 7 : 8. Hind femora with row of sparse black ventral

setae, longer than half of femora diameter, with one preapical

antero-dorsal seta. Hind tibia with ventral and dorsal rows of hairs,

as long as tibia diameter. Length ratio of hind coxa to femora to

tibia to tarsus (segments first to fifth) - 13 : 70 : 65 : 15 : 15 :

10 : 7 : 9.

Wings hyaline, veins brown. R1 ending at basal 1/3 of wing. Ratio

of parts of costa between R2+3 and R4+5 to those between R4+5 and M1 -

21 : 15. R4+5 and M1+2 straight, parallel in apical half. Ratio of

cross-vein m-cu to apical part of CuA1 - 13 : 20. Lower calipter

yellow, small, with short light hairs. Halters yellow.

Abdomen dark bronze-green, flattened dorso-ventrally,

subtriangular in dorsal view, with short dark setulae. Hypopygium

dark-brown. Cercus yellow, very long, band-like, approximately as long

as 4th and 5th terga combined, white setose along entire length, with

rounded apex; all hairs of equal length except the longest 2 or 3

apical cilia, 1/4 as long as cercus. Surstylus simple, narrow, thin in

apical third, ending at apical 1/3 length of cercus, with one strong

dorsal and one strong ventral preapical cilia, 2 apico-dorsal short

setulae.

Female. Similar to male except lacking male secondary sexual

characters; postero-dorsal seta on anterior tibia indiscernible.

Length: body 2.1 mm, antenna 0.4 mm, wing-length 2.3 mm,

wing-width 0.8 mm.

Diagnosis. This very small species can be recognized by 4

dorsocentral setae on mesonotum and very long male cercus reaching

almost to the base of abdomen. Legs yellow. Base of antenna yellow

ventrally. It can be hardly united with T. (Schoenophilus) versutus

Walker in one subgenus. Male from Angola (see description above)

slightly differs from the description of T. prudens by Curran and from

male collected in Senegal. Cercus of the last male somewhat longer, as

long as 3rd, 4th, and 5th terga, with about 10 apical cilia, nearly

half as long as cercus; surstylus ending in front of the middle of

cercus, with somewhat finer preapical cilia. Female from Senegal

differs from female collected in Angola only by distinctly developed

postero-dorsal seta on fore tibia.

Distribution. South Africa, Angola (!), Zaire, Ghana, Senegal (!).

Thinophilus imperialis (Curran)

Material examined. 7 males & 11 females, Gold Coast: Tamale,

November 1916, J.J. Simpson / Pres. by Imp. Inst. Ent. B.M. 1935-576.

Male, N. Nigeria: Ilorin, Apl. 1912, J.W. Scott Macfie, 1912-429

[NHML]. Male, N. Nigeria: Zaria, Samaru, 17.X.1968 / J.C. Deeming,

m.v. trap [NHML]. 2 females, Nigeria: Ile-Ife, W State, 10815 Aug.

1971, Col. J.T. Medler [NHML]. Female, Botswana (B24): R. Shashe, 20

mls. NW Francistown, 24.IV.1972 / Southern African Exp. B.M. 1972-1.

Diagnosis. T. imperialis differs from other species with long

curved apico-ventral setae on anterior tibia by very long cercus

extending to the base of abdomen.

Distribution. South Africa, Botswana (!), Congo, Ghana, Nigeria,

Zaire.

Thinophilus flavipalpis (Zetterstedt)

Material examined. 1 male & 2 females, Russia: Rostov Region,

Azov distr., Port-Katon, 3 & 9.VI.1996, Grichanov [Author's Coll.].

Male, North Kazakhstan: Tselinograd env., Novoishimka, 17.VII.1989,

Grichanov [Author's Coll.]. 30 males and females, Kirgizia: Issyk-Kul

Lake: Anan'evo, Kuturga, Darkhan, Ottuk, 1978-1979, Grichanov [VU]. 9

males and females, Ukraine: Odessa Region: Berezovka, 12.VIII.1977,

Tiligul'skii Liman, 11.VIII.1977, Bolgrad, 18.VIII.1977, Grichanov

[VU].

Diagnosis. This is a large species with black femora, ornamented

anterior tarsus, relatively simple middle tarsus, and short cercus. It

is closely related to T. ornatus Negrobov et Grichanov.

Distribution. Egypt, West Europe (except North), Ukraine (Odessa

Region, Zaporozh'e Region, Crimea), South Russia (Rostov Region),

Caucasus, Kazakhstan, Kirgizia, Mongolia; ?Mozambique.

Thinophilus brevicilius Negrobov

Material examined. 2 males and 1 female, Kirgizia: Issyk-Kul

Lake: Anan'evo, Ottuk, 1978-1979, Grichanov [VU].

Diagnosis. T. brevicilius is closely related to T. setosus

Negrobov and similar to other species having black femora and

ornamented anterior tarsus, differing from these by simple 3rd and 4th

tarsomeres of anterior tarsus, posterior femora without long ventral

setae, and mostly yellow tibiae.

Distribution. Uzbekistan, Tadjikistan, Kirgizia (!).

Thinophilus mirandus Becker

(Fig. 1)

Material examined. 3 males and 2 females, Tanzania: Matombo,

Morogoro reg. / 11.IX.1977, leg. Mahunka [HNHM].

Description. Frons bronze-green, violet in the middle, slightly

grey-brownish pollinose. A row of black postocular setae ending with

postvertical seta at the top of eye present. One black vertical seta

laterally on frons. Ocellar tubercle with one pair of strong setae and

several hairs. Ventral postcranium covered with dense long white

irregular setae. Face metallic blue-green, slightly pollinose, the

narrowest under antennae. Ratio of height of epistome to its minimal

width to height of clypeus to its maximal width to height of palpus,

21 : 12 : 10 : 20 : 30. Antenna short, entirely yellow-orange, first

flagellomere rounded, with short terminal hairs. Arista dorsal, bare,

brownish, thick at base. Length ratio of scape to pedicel to first

flagellomere to arista - 5 : 5 : 9 : 31. Palpus yellow, white

pollinose, with sparse long black hairs. Proboscis brown, with white

hairs.

Thorax olive-bronze-green, entirely grey-brownish pollinose,

without dark spots. Six dorsocentral bristles, the last one is the

longest; no acrostichals. Prothorax and propleura with a few white

hairs. Scutellum with 2 strong black setae and 2 very small hairs.

Legs yellow, apical segments of tarsi partly darkened. Fore coxa

yellow, white pollinose, with black hairs. Middle and hind coxae

olive-green except apex, grey pollinose, each with one external seta,

middle coxa also with dark hairs. Fore femora with a row of black

poster-ventral hairs in apical half, half as long as diameter of

femora. Fore tibia with 2 rows of somewhat elongated ventral setulae,

and a row of dorsal setae of various length, with 3 long black curved

preapical postero-ventral setae; antero-ventral side bare. First

tarsomere with nearly right angle bend, with ventral excavation at

base; baso-ventral prominence bearing a group of very short spinules;

1st and 2nd tarsomeres with a row of elongated posterior setulae; 3rd

and 4th tarsomeres each with a group of long black dorsal hairs, twice

as long as diameter of tarsomere; 4th tarsomere also with one long

dorsal seta, as long as three last segments of fore tarsus together;

5th tarsomere whitish and flattened; all tarsomeres bare ventrally and

darkened at tip. Length ratio of fore coxa to femora to tibia to

tarsus (segments from first to fifth) - 30 : 48 : 40 : 19 : 9 : 6 : 5

: 8. Middle femora with a few dark fine ventral setae in apical two

thirds, nearly half as long as diameter of femora, and with a few

stronger black posterior preapical setae. Middle tibia with two or

three antero-dorsal, two or three postero-dorsal and one ventral

setae. Middle tarsomeres each with a few strong apical setulae; 4th

and 5th tarsomeres flattened, partly white, ventrally bare. Length

ratio of middle coxa to femora to tibia to tarsus (segments from first

to fifth) - 26 : 53 : 57 : 29 : 10 : 7 : 6 : 10. Hind femora with

poster-dorsal and postero-ventral hairs in apical half, shorter than

half-diameter of femora. Hind tibia with 4 or 5 setae on each of

antero-dorsal, postero-dorsal, antero-ventral and postero-ventral

sides. 5th tarsomere slightly flattened, ventrally bare. Length ratio

of hind coxa to femora to tibia to tarsus (segments first to fifth) -

18 : 66 : 75 : 19 : 15 : 10 : 7 : 9.

Wings hyaline, veins mostly brown, partly yellow in basal half.

R1 short, extending to basal third of wing. Ratio of parts of costa

between R2+3 and R4+5 to those between R4+5 and M1 - 26 : 16. R4+5 and

M1+2 nearly parallel at apex, without distinct curvations. Ratio of

cross-vein m-cu to apical part of CuA1 - 20 : 35. Lower calipter

yellow, with white hairs. Halters yellow.

Abdomen olive-bronze-green, with short dark setae; 6th segment

darker; 8th segment with white pubescence and dark hairs. Hypopygium

dark-brown, white pollinose. Cercus yellow, band-like, with rounded

apex, somehow protruding behind the end of surstylus. Surstylus

brown, stick-shape, densely setose in apical third, with two dorsal

setae in the middle. Two epandrial setae at base of surstylus present.

Hypandrium assymmetric, nearly as wide as cercus, reaching the end of

surstylus.

Female. Similar to male except lacking male secondary sexual

characters, otherwise as follows: ratio of height of epistome to its

minimal width to height of clypeus to its maximal width to height of

palpus, 21 : 17 : 13 : 25 : 30. Femora lacking long setae except

several long postero-dorsal hairs in apical third of hind femora. Fore

tibia with 3 strong antero-dorsal, 3 short postero-dorsal and 3 short

ventral setae. Tarsi simple, but 5th tarsomere of all tarsi slightly

flattened.

Length: body without antennae 3.9 - 5.5 mm, wing-length 3.9 - 5.3

mm, wing-width 1.3 - 1.6 mm.

Diagnosis. T. mirandus is easily recognized by the anterior

basitarsomere having nearly right angle bend. Middle femora with

postero-ventral setae in middle part, at least half as long as femora

diameter. Females can be separated by 6 dorsocentrals, entirely yellow

antenna, black setation on palpus, and maculated wing.

Distribution: Algeria, Morocco, Tanzania (!).

Thinophilus spinitarsis Becker

Material examined. Male, South Tadjikistan: Dusti env.,

13.VII.1991, Grichanov [Author's Coll.].

Diagnosis. T. spinitarsis is closely related to T.

vanschuytbroecki Negrobov. It differs from other species with yellow

femora and ornamented anterior tarsus by fairly curved anterior

basitarsomere, hyaline wing, and long dorsal lobe of surstylus in male

genitalia.

Distribution. China, Iran, Tadjikistan (!), South Ukraine

(Kherson Region).

Thinophilus munroi munroi Curran

Material examined. Female, S. Africa: R.E. Turner, Brit. Mus.

1935-73 / Cape Province, Mossel Bay, Dec. 1934.

Diagnosis. T. munroi munroi strongly differs from all other

species by setation of scutellum having 6 to 8 pairs of short spines

in addition to one pair of long setae. It is closely related to T.

munroi setiscutellatus ssp.n., differing by distinctly annulate

posterior tarsus in both sexes, dark setation on anterior coxa and

femora, and many other characters. Wing vein R1 conspicuously shorter

than in T. munroi setiscutellatus, being also unusually long.

Distribution. South Africa.

Thinophilus munroi setiscutellatus ssp.n.

(Fig. 2)

Holotype, male; paratypes, 7 males and 4 females, S.W. Africa

(25), Swakopmund, 26-30.I.1972 / Southern African Exp. B.M. 1972-1.

Description. Frons and occiput bronze-green, densely

grey-brownish pollinose. 7 to 10 short black postocular setae in

irregular upper row on either side, the last upper seta slightly

stronger. One short black vertical seta laterally on frons. One pair

of ocellar setae. Ventral postcranium covered with dense dirty-white

irregular postocular setae. Face densely grey-yellowish pollinose, the

narrowest under antennae. Ratio of height of epistome to its minimal

width to height of clypeus to its maximal width to height of palpus,

20 : 17 : 11 : 24 : 15. Terminal margin of clypeus covered with short

dense white hairs. Antenna short, orange, first flagellomere brown in

apical third, rounded, with short terminal hairs. Arista dorsal, bare,

thick and brown at base, otherwise fine and whitish. Length ratio of

scape to pedicel to first flagellomere to arista - 4 : 3 : 9 : 35.

Palpus whitish-yellow, greyish-white pollinose, with sparse long white

hairs. Proboscis black, with white hairs.

Thorax olive-bronze-green, entirely grey-brownish pollinose,

without dark spots. Six or seven dorsocentral bristles, the last one

is the longest; no acrostichals. Propleura with a few white hairs.

Scutellum with 3 to 5 short black setae on each side in addition to a

pair of normal long bristles.

Legs yellow, apical segments of fore and middle tarsi and apices

of all tarsomeres of hind tarsus darkened. Fore coxa olive-green,

entirely white pollinose, with short white hairs. Middle and hind

coxae olive-green except apex, grey pollinose, with a few short dark

hairs. Fore femora with a few light ventral hairs at base, half as

long as diameter of femora. Fore tibia with very small, almost

indiscernible setae: two antero-dorsal, two postero-dorsal and one

ventral. Length ratio of fore coxa to femora to tibia to tarsus

(segments from first to fifth) - 30 : 47 : 46 : 19 : 8 : 7 : 5 : 7.

Middle femora bears a group of dark antero-ventral hairs in apical

half, hardly as long as diameter of femora, also with one stronger

preapical anterior seta of the same length, and with a few very short

fine white hairs in basal half. Middle tibia with very small setae:

two antero-dorsal, two postero-dorsal and one postero-ventral. Length

ratio of middle coxa to femora to tibia to tarsus (segments from first

to fifth) - 22 : 54 : 57 : 30 : 10 : 8 : 5 : 7. Hind femora without

long hairs. Hind tibia with short setae: two antero-dorsal, two

postero-dorsal and one ventral. Hind basitarsomere slightly broadened

at apex, with apical group of short dense posterior setulae. Length

ratio of hind coxa to femora to tibia to tarsus (segments first to

fifth) - 15 : 80 : 66 : 25 : 14 : 10 : 6 : 9.

Wings hyaline, veins mostly brown, partly yellow in basal half.

R1 long, extending to the middle of wing. Ratio of parts of costa

between R2+3 and R4+5 to those between R4+5 and M1 - 20 : 14. R4+5 and

M1+2 nearly parallel at apex. M1+2 with indistinct curvation. Ratio of

cross-vein m-cu to apical part of CuA1 - 22 : 47. Lower calipter

yellow, with short yellow hairs. Halters yellow.

Abdomen olive-bronze-green, with short dark setulae. Hypopygium

dark-brown, with white pubescence. Cercus yellow, densely white

setose along entire length, spatulate, flat, band-like with rounded

apex, narrowed distad; all hairs of equal length, approximately as

long as width of cercus. Surstylus brown, bilobate, with thin lobes,

approximately as long as cercus, although protruding behind the end of

cercus; external lobe shorter, flat, with 3 apical and 1 subapical

spinose setae; interior lobe longer, cylindrical, narrowed apicad, with

1 dorsal seta in the middle, 1 apical spinose seta, and several

subapical hairs. Epandrial lobe at base of surstylus present. having

long apical seta. Hypandrium broadened in distal half, as wide as

cercus, narrowed at apex, pointed, reaching the end of external lobe

of surstylus.

Female. Similar to male except lacking male secondary sexual

characters, otherwise as follows: ratio of height of epistome to its

minimal width to height of clypeus to its maximal width to height of

palpus, 25 : 21 : 13 : 29 : 20. Arista whitish. Palpus with short

white hairs. Upper row of 6 or 7 short black postocular setae present,

separate row of 2 or 3 same setae at the top of eye, one short

occipital seta, one vertical seta. Fore and middle femora without long

hairs. Otherwise similar to male.

Length: male body without antennae 4.5 - 4.7 mm, wing-length 4.3

- 5.2 mm, wing-width 1.4 - 1.6 mm.

Distribution: Namibia.

Etymology. The subspecies name reflects the presence of

excessively developed setation on scutellum.

Diagnosis. T. munroi setiscutellatus is an allied subspecies for

T. munroi munroi. It can be separated by weaker developed setation on

scutellum, only pale hairs on anterior coxa and femora, and not so

strongly annulate posterior tarsus as in T. munroi munroi. Females

have the same differences. The species forms a separate group,

possibly, subgenus, having such characters as long vein R1 and

excessively setose scutellum.

Thinophilus ciliventris sp.n.

(Fig. 3)

Holotype, male, S. Africa (S19): R. Magalakwena, 23o26' S, 28o37'

E, 26.IV.1972 / Southern African Exp. B.M. 1972-1.

Paratypes, 2 females, the same labels; 1 male & 3 females, Angola

(A32): Cachoeiras, 20 mls. SW Gabela, 18-19.III.1972 / Southern

African Exp. B.M. 1972-1; 2 females, Botswana (B24): R. Shashe, 20

mls. NW Francistown, 24.IV.1972 / Southern African Exp. B.M. 1972-1;

male, N. Nigeria: Zaria, Samaru, 1.VII.1969 / J.C. Deeming, m.v. trap

[NHML].

Description. Frons blue-violet, slightly pollinose. A row of

short black postocular setae ending with a few postvertical hairs at

the top of eye present; one strong occipital seta; one black vertical

seta laterally on frons. Ocellar tubercle with one pair of strong

setae and several short hairs. Ventral postcranium covered with long

dense white irregular setae. Face violet, slightly pollinose, the

narrowest in the middle of epistome. Ratio of height of epistome to

its minimal width to height of clypeus to its maximal width to height

of palpus, 26 : 17 : 14 : 25 : 33. Antenna short, mostly dark-brown,

all segments ventrally yellow-orange, scape with pale interior spine,

first flagellomere rounded, with short terminal hairs. Arista dorsal,

bare, brown, thick at base, otherwise fine. Length ratio of scape to

pedicel to first flagellomere to arista, 12 : 9 : 13 : 51. Palpus

yellow, white pollinose, with sparse short black hairs. Proboscis

black, with white hairs.

Mesonotum metallic green with copper and blue reflection mostly

grey-brownish pollinose, without dark spots. Pleura olive-green, grey

pollinose. Six or seven dorsocentral bristles, the last one is the

longest; no acrostichals. Propleura and prothorax with long white

hairs. Scutellum with 2 strong black setae and 2 very short hairs.

Legs mostly yellow-orange, apical segments of fore and middle

tarsi darkened, apex of hind femora, extreme base and apex of hind

tibia, and hind tarsus brown. Fore coxa olive-green at base, entirely

white pollinose, with dark hairs in apical half, with shorter white

hairs in basal half. Middle and hind coxae olive-green except apex,

grey pollinose, each with one external seta; middle coxa also with a

few short dark hairs, hind coxa with microscopic light hairs. Fore

femora with a row of black ventral setae, half as long as diameter of

femora, with a few longer poster-ventral setae in apical fourth. Fore

tibia with 3 to 5 dorsal and several simple apical setae, with a row

of ventral setulae, more than half as long as tibia diameter, and a

row of shorter antero-ventral spinules along entire length; anterior

side bare. First tarsomere with ventral row of spinules, half as long

as diameter of tarsomere; other tarsomeres with somewhat elongated

terminal setulae. Length ratio of fore coxa to femora to tibia to

tarsus (segments from first to fifth) - 36 : 60 : 50 : 17 : 8 : 6 : 5

: 7. Middle femora with very short black ventral setae in basal half,

also with one stronger preapical anterior seta. Middle tibia with

three antero-dorsal, two postero-dorsal, one antero-ventral and three

postero-ventral setae of various strength. Last three tarsomeres

slightly flattened, with bare ventral side. Length ratio of middle

coxa to femora to tibia to tarsus (segments from first to fifth) - 29

: 63 : 73 : 42 : 13 : 8 : 6 : 9. Hind femora with ventral and

antero-dorsal rows of seta, shorter than half-diameter of femora. Hind

tibia with four antero-dorsal, four postero-dorsal and three ventral

setae of various strength. Length ratio of hind coxa to femora to

tibia to tarsus (segments first to fifth) - 19 : 95 : 105 : 26 : 19 :

11 : 8 : 9.

Wings practically hyaline, veins mostly brown, partly yellow at

base. R1 short, extending to basal third of wing. Ratio of parts of

costa between R2+3 and R4+5 to those between R4+5 and M1 - 30 : 20.

R4+5 and M1+2 nearly parallel at apex, M1+2 without distinct

curvations. Ratio of cross-vein m-cu to apical part of CuA1 - 27 : 29.

Lower calipter yellow, with short white hairs. Halters yellow.

Abdomen mostly bronze-green with copper reflection, with short

dark setulae; 5th segment with violet reflection; 6th segment entirely

violet; 3rd and 4th sternites with a tuft of numerous long white hairs;

8th segment with white pollination and white pubescence. Hypopygium

black, white pollinose. Cercus light-brown, long, flat, band-like,

pointed on apex, with short dark hairs along entire length and one

somewhat longer cilia on extreme apex, reaching the end of surstylus.

Surstylus brown, broad and flat, mitten-like, with dorso-lateral

horn-like process bearing two small setae, with two cilia near

epandrium, and several scattered short hairs. Hypandrium narrow,

slightly broadened in distal half, reaching the middle of surstylus.

Female. Similar to male except lacking male secondary sexual

characters, otherwise as follows: anterior coxa sometimes imparting

mostly olive-green; fore femora without strong setae; anterior tibia

with 3 or 4 antero-dorsal, 3 or 4 postero-dorsal and 2 or 3 ventral

setae; tarsi simple.

Length: male body without antennae 5.8 - 7.1 mm, wing-length 4.8

- 6.2 mm, wing-width 1.4 - 1.6 mm; female body 5.6 - 6.1 mm, wing 5.5

- 6.0 mm.

Distribution: South Africa, Botswana, Angola, Nigeria.

Etymology. The species name reflects the presence of long cilia

on abdominal venter.

Diagnosis. Judging from the examined material and published

descriptions, none Afrotropical or Palearctic species has abdominal

venter with long white hairs. Only T. bicalcaratus bears 2 bundles of

strong black bristles on the 4th sternite of abdomen. Other diagnostic

characters are as follows: anterior coxa yellow haired in basal half;

posterior femora brown from above in apical third, without long

antero-ventral setae; fore tibia and basitarsomere with ventral row of

very short spinules along entire length; propleura with white

setation; m-cu as long as apical part of CuA1.

Thinophilus palpatus Parent

(Fig. 4)

Material examined. Male, N. Nigeria: Zaria, Samaru, 23.II.1968 /

J.C. Deeming, m.v. trap [NHML]; female, Nigeria: N.W. State, Mokwa,

I.A.R., Mile 1, 8-17.VIII.1970, P.H. Ward, B.M. 1970-604; 2 females,

N. Nigeria: Ilorin, 22.02.1912, Dr. J.V. Scott-Macfie / Pres. by Imp.

Inst. Ent. Brit. Mus. 1931-287; male, S.W. Africa (W1): Nr.

Onseepkans, Orange R. banks, 8-10.I.1972 / Southern African Exp. B.M.

1972-1; female, S.W. Africa: Otjitambi Fm., 27 mls ESE Kamanjab,

13-15.II.1972 / Southern African Exp. B.M. 1972-1; male, S.W. Africa

(W30): Ameib Farm, 31.I-2.II.1972, on vegetation around pools /

Southern African Exp. B.M. 1972-1; 2 females, Angola (A2): Rocadas, R.

Cumene, 19-22.II.1972 / at light / Southern African Exp. B.M. 1972-1;

8 females, Angola (A32): Cachoeiras, 20 mls. SW Gabela, 18-19.III.1972

/ Southern African Exp. B.M. 1972-1; 2 females, Angola (A16): 2 mls.

N. Mocamedes, 29.II.1972 / Southern African Exp. B.M. 1972-1; female,

Angola (A37): 5 mls. NE Negola, 25.III.1972 / Southern African Exp.

B.M. 1972-1; female, Botswana (B7): Kuke Pan, 20o59'S, 22o25'E,

14-15.IV.1972 / Southern African Exp. B.M. 1972-1; female, Botswana

(B16): Maun, 21.IV.1972 / Southern African Exp. B.M. 1972-1;

Diagnosis. This is a comparatively small (3.5 mm) species with

simple yellow legs, narrow face, leaf-like male cercus, differing from

T. ruficornis and related species mainly by hypopygium morphology and

wing veins CuA1/m-cu length ratio.

Distribution. Ethiopia, Nigeria, Angola (!), Namibia (!),

Botswana (!).

Thinophilus ruficornis (Haliday)

Material examined. 3 males & 3 females, South Russia: Rostov

Region, Azov distr., Port-Katon, 3, 9, 18 & 26.06.1996, Grichanov

[Author's Coll.]. 7 males & females, Ukraine: Odessa Region:

Khadzhibeiskii Liman, 28.VIII.1977, Tiligulskii Liman, 19.VII.1976,

Berezovka, 12.VIII.1977, Grichanov [VU]. Male, Russia: Novosibirsk

Region, Krasnozerskoe, 20.VI.1989, Grichanov [Author's Coll.]. 12

males & females, Kirgizia: Issyk-Kul Lake: Anan'evo, Kuturga, Ottuk;

1978-1979, Grichanov [VU].

Diagnosis. This is a comparatively small species with simple

yellow legs, narrow face, short and narrow male cercus, and small

surstylus.

Distribution. Whole Europe, West Siberia (Omsk Region,

Novosibirsk Region), North Kazakhstan, Kirgizia, Mongolia, North

China.

Thinophilus argyropalpis Becker

Material examined. 2 males, South Tadjikistan: Dusti env.,

22 & 24.VI.1985, Grichanov & Shamshev [Author's Coll.].

Diagnosis. T. argyropalpis is small species with simple yellow

legs, closely related to T. vanschuytbroecki Negrobov. It differs from

other species by only two setae on scutellum, silvery-white palpus,

grey pollinose face, and partly yellow antenna.

Distribution. Egypt, Tunisia, South Arabia, Iran, Uzbekistan,

Turkmenia, Tadjikistan (!), Kazakhstan, South Russia (Volgograd

Region), Ukraine (Odessa Region).
 
 
 
 
 
 

Key to Afrotropical and Palearctic species of Thinophilus

1. Mesonotum and/or scutellum with distinct dark lateral spots.......2

- Mesonotum without dark lateral spots..............................8

2. Wing with dark spot near the end of R2+3 and R4+5; anterior spot of

mesonotum nearly as large as notopleura, no prescutellar spot......

.............................................quadrimaculatus Becker

- No spot at wing apex..............................................3

3. Mesonotum with additional spot in front of scutellum..............4

- Mesonotum without spot in front of scutellum......................6

4. Mesonotum with four lateral spots..................indigenus Becker

- Mesonotum with six lateral spots...................................5

5. Anterior basitarsomere with simple setulae, without spinules; small

species (2.75 mm)..................................maculatus Parent

- Anterior basitarsomere with a row of very short spinules...........

.........................................splendidus Vanschuytbroeck

6. Wing distinctly maculated at m-cu and on M1+2 curvation............

.................................................bipunctatus Curran

- Wing hyaline......................................................7

7. 5 dorsocentrals; scutellum with one spot; legs mostly yellow,

anterior femora without ventral comb of setae, middle tibia curved

in the middle....................................setulipalpis Bezzi

- At least 6 dorsocentrals; scutellum with 2 lateral spots; legs

mostly black, anterior femora with ventral comb of black setae,

middle tibia simple......................................rex Curran

8. 4 dorsocentrals; small species (2 - 2.5 mm).......................9

- 5 or 6 dorsocentrals present, front one usually short; size usually

larger than 3 mm.................................................10

9. All male femora with long ventral setae; cercus long, reaching

almost to the base of abdomen; legs yellow...........prudens Curran

- Only middle femora with a row of ventral setae, not longer

than femora diameter; cercus very short, triangular-oval; legs

brownish or greyish-yellow with yellow knees.......................

....................................(Schoenophilus) versutus Walker

10. Pedicel long, with apico-ventral lobe..............promotus Becker

- Pedicel without apico-ventral lobe..............................11

11. Scutellum with 3 to 8 pairs of short spines in addition to one

pair of long setae; 1st and 2nd tarsomeres of hind tarsus annulate

with yellow and black............................................12

- Scutellum with at most two pairs of setae........................13

12. Anterior coxa with black setae and pale hairs; anterior femora

with a few long black ventral hairs in basal third; scutellum with

6 to 8 pairs of spines.........................munroi munroi Curran

- Anterior coxa and femora with pale hairs only; scutellum with 3 to

5 pairs of spines.....................munroi setiscutellatus ssp.n.

13. Anterior four femora at least partly black (females usually

indeterminable); male anterior tibia with 2 or 3 strong curved

apical postero-ventral setae.....................................14

- Anterior four femora yellow, sometimes infuscated from above;

anterior tibia with or without apical setae......................18

14. Anterior femora black from above, other femora mostly

reddish-yellow; male cercus very long, extending to the base of

abdomen........................................imperialis (Curran)

- All femora partly black; cercus much shorter....................15

15. 2nd, 3rd and 4th tarsomeres of anterior tarsus with a group of

black setae, that longer than article diameter...................16

- 3rd and 4th tarsomeres of anterior tarsus without long setae.....17

16. 2nd and 3rd tarsomeres of middle tarsus with long flattened setae.

......................................ornatus Negrobov et Grichanov

- Middle tarsus with simple setae...........flavipalpis (Zetterstedt)

17. Posterior femora with ventral setae, half as long as femora

diameter; tibiae mostly dark.......................setosus Negrobov

- Posterior femora ventrally with short setulae; tibiae mostly yellow

...............................................brevicilius Negrobov

18. Males...........................................................19

- Females.........................................................38

19. Posterior coxa with curved apical internal spine; 5 dorsocentrals

present; scutellum with 2 setae..................ochripalpis Becker

- Posterior coxa without spine.....................................20

20. Anterior basitarsomere with internal excavation or incision at

base; antenna yellow.............................................21

- Anterior basitarsomere without excavation or incision at base....23

21. Anterior basitarsomere with nearly right angle bend; middle femora

with postero-ventral setae in middle part, at least half

as long as femora diameter..........................mirandus Becker

- Anterior basitarsomere fairly curved; middle femora with short

setulae in middle part...........................................22

22. Dorsal lobe of surstylus short; wing near m-cu and on M1+2

curvation distinctly maculated; anterior coxa dark in basal

part......................................vanschuytbroecki Negrobov

- Dorsal lobe of surstylus long; wing practically hyaline; anterior

coxa with dark spot near base....................spinitarsis Becker

23. Anterior and/or middle femora ventrally with hairs and bristles,

nearly as long as femora diameter...............................24

- Anterior and middle femora without long ciliation...............31

24. Palpus with black hairs.........................................25

- Palpus with white hairs.........................................29

25. Anterior basitarsomere with ventral row of short but strong black

spines, at least half as long as article diameter................26

- Anterior basitarsomere without ventral spines, with simple setulae

only.............................................................27

26. Apical half of apical part of M1+2 distinctly arcuate, m-cu 2/3 as

long as apical part of CuA1, length 4 mm..........spinulosus Parent

- M1+2 evenly curved, m-cu as long as apical part of CuA1, length 6

to 7 mm...........................................ciliventris sp.n.

27. Wing with spots at m-cu and on M1+2 curvation; mesonotum with a

black spot in front of scutellum; m-cu as long as apical part of

CuA1, length 2.75 mm...............................maculatus Parent

- Wing and mesonotum without such spots; length 3.5 mm.............28

28. Cercus gradually pointed towards apex; m-cu 2/3 as long as apical

part of CuA1........................................capensis Curran

- Cercus broad in the middle, narrowed towards apex; m-cu as long

as apical part of CuA1..............................palpatus Parent

29. Anterior coxa yellow...........................longipilus Negrobov

- Anterior coxa black.............................................30

30. 5 dorsocentrals present; last tarsomeres black on apex; arista

black; size 4.5 mm.....................................calopus Loew

- 6 or 7 dorsocentrals present; last tarsomeres entirely black;

arista white; size 6 mm.............................virgatus Curran

31. Scutellum with 2 strong and 2 short setae; face nearly twice as

high as wide near suture.........................................32

- Scutellum with 2 setae; face approximately as high as wide near

suture...........................................................36

32. 4th sternite of abdomen with 2 bundles of strong black bristles;

anterior coxa with black hairs; posterior femora with long

antero-ventral setae near apex...............bicalcaratrus Negrobov

- 4th sternite of abdomen without bundles of black bristles; anterior

coxa yellow haired in basal half; posterior femora without long

antero-ventral setae.............................................33

33. 3rd and 4th sternites with a tuft of long white hairs; fore tibia

and basitarsomere with ventral row of very short spinules along

entire length.....................................ciliventris sp.n.

- Abdominal venter without tuft of long hairs......................34

34. Anterior basitarsomere with postero-ventral setae longer than

tarsomere diameter; apical part of CuA1 thrice as long as m-cu.....

....................................................pollinosus Loew

- Anterior basitarsomere without long setae; apical part of CuA1 no

more than twice as long as m-cu..................................35

35. Cercus short and narrow, shorter than surstylus; surstylus small,

dorsal lobe hook-shaped on apex, without long dorsal setae; apical

part of CuA1 approximately twice as long as m-cu...................

...............................................ruficornis (Haliday)

- Cercus broad, leaf-shaped, longer than surstylus; surstylus

narrowed towards apex; apical part of CuA1 approximately as long as

m-cu.................................................palpatus Parent

36. Face shining metallic, slightly pollinose; sutural setae

distinctly developed; abdomen with long hairs......achylleus Becker

- Face silvery or greyish-white pollinose; sutural setae small, 1/3 to

1/4 as long as supraalary setae; abdomen with short hairs........37

37. Palpus yellow; face silvery-white pollinose; antenna mostly dark...

..........................................vanschuytbroecki Negrobov

- Palpus silvery-white; face grey pollinose; antenna distinctly

yellow from below...............................argyropalpis Becker

38. 5 dorsocentrals.................................................39

- At least 6 dorsocentrals present, front one usually short........44

39. Palpus with white hairs.........................................40

- Palpus with black hairs.........................................41

40. Anterior coxa yellow............................quadrisetus Parent

- All coxae black.......................................calopus Loew

41. Antenna entirely yellow.........................ochripalpis Becker

- Antenna partly black............................................42

42. Wing strongly infuscated............................tinctus Parent

- Wing hyaline, at most with spots at m-cu and on M1+2 curvation...43

43. Tarsi entirely black.............................atritarsis Parent

- Tarsi partly light................................aquaticus Becker

44. Antenna entirely yellow.........................................45

- Antenna partly black............................................50

45. Palpus with pale setation.......................................46

- Palpus with black setation......................................48

46. Face shining metallic, practically without pollination; scutellum

with 2 setae.........................................achylleus Mik

- Face pollinose; if face weakly pollinose, than scutellum with 2

strong and 2 small setae........................................47

47. Anterior coxa yellow............................quadrisetus Parent

- All coxae black.......................................calopus Loew

48. Wing without spots; tarsi at least partly black...spinulosus Parent

- Wing maculated with 2 smoky spots on m-cu and M1+2 curvation....49

49. Scutellum with 2 setae; 5 dorsocentrals.........ochripalpis Becker

- Scutellum with 2 strong and 2 small setae; 6 dorsocentrals........

...................................................mirandus Becker

50. Palpus with pale setation.......................................51

- Palpus with black setation......................................55

51. Scutellum with 4 setae..........................quadrisetus Parent

- Scutellum with 2 setae..........................................52

52. Hind tarsomeres annulate with yellow and black; 8 dorsocentrals;

no distinct prothoracic setulae; anterior coxa with exclusively

pale fine hairs................................annulitarsis Parent

- Hind tarsomeres non-annulate....................................53

53. Palpus silvery-white; tarsi black; anterior coxa with pale hairs..

...............................................argyropalpis Becker

- Palpus yellow-orange; tarsi mostly yellow.......................54

54. Anterior coxa with pale hairs............vanschuytbroecki Negrobov

- Anterior coxa with black hairs.....................virgatus Curran

55. Hind femora with long dorsal setae..............................56

- Hind femora without long dorsal setation, at most with single

anterior preapical seta.........................................59

56. Upper postocular setae in two rows.................modestus Becker

- Upper postocular setae in one row...............................57

57. Propleura with 2 or 3 strong black setae and white hairs; hind

femora yellow..................................longipilus Negrobov

- Propleura with white setation...................................58

58. Hind femora brown from above in apical third.....ciliventris sp.n.

- Hind femora entirely yellow........................palpatus Parent

59. Apical part of CuA1 thrice as long as m-cu; posterior coxa mostly

yellow.............................................pollinosus Loew

- Apical part of CuA1 no more than twice as long as m-cu..........60

60. Apical part of CuA1 1/3 longer than m-cu; posterior coxa mostly

dark; tibiae at base and at apex dark..............capensis Curran

- Apical part of CuA1 approximately twice as long as m-cu; posterior

coxa mostly yellow; all tarsi and apex of tibiae dark.............

..............................................ruficornis (Haliday)

Acknowledgements

I am sincerely grateful to Dr. Brian Pitkin, Dr. Laslo Papp, and

Dr. Roy Danielsson for their kindness in furnishing an opportunity to

study the collections of the Natural History Museum (London), the

Hungarian Natural History Museum (Budapest), and Lund University.

References

Curran C.H. 1926. Records of African Dolichopodidae with

descriptions of new species. Rev. Zool. Afr., 14(1): 1-39.

Dyte C.E. & Smith K.G.V. 1980. Family Dolichopodidae. In R.W.

Crosskey (ed.). Catalogue of the Diptera of the Afrotropical Region.

Brit. Mus. (Nat. Hist.), London: 443-463.

Grootaert, P. & Meuffels, H.J.G. 1984. Dolichopodidae (Diptera)

from Papua New Guinea. II. Some new species of the genus Thinophilus

Wahlberg, 1844, from the lowland. Indo-Malayan Zoology, 2: 209-223.

Negrobov O.P. 1991. Dolichopodidae. In Catalogue of Palaearctic

Diptera. V.7. Dolichopodidae-Platypezidae. Akad. kiado. Budapest:

1-291.

Remark under figures

Figs 1-4. Surstylus (=gonopode), lateral view.

1, T. mirandus Becker;

2, T. munroi setiscutellatus ssp.n.;

3, T. ciliventris sp.n.;

4, T. palpatus Parent.

Fig. 5, T. prudens Curran, hypopygium, lateral view.