CLASSIFICATION OF DOLICHOPODOIDAE
by Igor Grichanov
Last updated: 4 December 2024I follow here Yang et al. (2006) mainly, referring all known genera to the described subfamilies despite critical notes by Sinclair et al. (2008). For example, Sinclair et al. wrote that the genera of the subfamilies Antyxinae and Kowmunginae would have been better placed as incertae sedis in the Dolichopodidae ''until a more rigorous phylogenetic study determines their placement as well as the status of the other subfamilies''. Many genera were recently described or declared as incertae sedis in the Dolichopodidae. I am afraid that their uncertain position bears a risk to leave them outside future revisions of subfamilies on a global scale. Therefore, I associate all those genera with one of 18 subfamilies of Dolichopodidae (including Parathalassiinae) based on Yang et al. (2006) with some changes based on the most recent literature and on my experience of treating dolichopodid genera. In addition, I divide here large subfamilies into tribes (if they were once proposed). None new taxon is here described. The sign † ahead of a name means extinct taxon.
EPIFAMILY DOLICHOPODOIDAE (=DOLICHOPODIDAE SENSU LATO)
FAMILY MICROPHORIDAE Collin, 1960
† Archichrysotus Negrobov, 1978:84
† Avenaphora Grimaldi et Cumming, 1999: 61
† Curvus Kaddumi, 2005: 85
† Electrophorella Cumming et Brooks, 2002: 42
† Meghyperiella Meunier, 1908: 112
† Microphorites Hennig, 1971:16
Microphor Macquart, 1827: 139
Schistostoma Becker, 1902:46
FAMILY DOLICHOPODIDAE Latreille, 1809
† Subfamily Eodolichopoditinae Hong, 2002
Remarks. Eodolichopoditinae is probably an unavailable name, because
the genus-group name Eodolichopodites is based on an unavailable species-group
names proposed after 1999 without specification of a type depository (Hong,
2002). The latter belongs also to the genera Bicercites, Columnocorna,
Eodolichopodites, Haodolichopodites, Laticopulus, Longilabia, Paradolichopodites,
Sinodolichopodites, Sunodolichopodites and Wangia de-scribed by the same
author. Hong mentioned 2001 as the year of description of his new taxa
everywhere in his big work (2002); however, I have not found his work with
the new taxa descriptions published in 2001.
† Bicercites Hong, 2002: 392
† Columnocorna Hong, 2002: 388
† Eodolichopodites Hong, 2002: 360
† Haodolichopodites Hong, 2002: 369
† Laticopulus Hong, 2002: 380
† Longilabia Hong, 2002: 373
† Orbilabia Hong, 2002: 368
† Paradolichopodites Hong, 2002: 382
† Septocellula Hong, 1981: 109
† Sinodolichopodites Hong, 2002: 377
† Sunodolichopodites Hong, 2002: 385
† Wangia Hong, 2002: 354
Subfamily Parathalassiinae Chvala, 1981
† Cretomicrophorus Negrobov, 1978:86
† Retinitus Negrobov, 1978:86
Amphithalassius Ulrich, 1991: 200
Chimerothalassius Shamshev et Grootaert, 2003:130
Eothalassius Shamshev et Grootaert, 2005:108
Microphorella Becker, 1909:28
Neothalassius Brooks et Cumming, 2016
Parathalassius Mik, 1891:217
Plesiothalassius Ulrich,1991: 191
Thalassophorus Saigusa,1986:106
Subfamily Achalcinae Grootaert et Meuffels, 1997
Achalcus Loew, 1857:30
Apterachalcus Bickel, 1991:12
Australachalcus Pollet, 2005:33
Scepastopyga Grootaert et Meuffels, 1997: 1588
Xanthina Aldrich, 1902: 92
Subfamily Antyxinae Yang, Zhu, Wang et Zhang, 2006
Antyx Meuffels et Grootaert, 1991: 290
Subfamily Babindellinae Bickel, 1987
Babindella Bickel, 1987: 97
Subfamily Diaphorinae Schiner, 1864
† Gheynia Meunier, 1899: 322
† Palaeoargyra Meunier, 1895: 5
† Palaeomedeterus Meunier, 1894: 121
† Prochrysotus Meunier, 1908: 9
Acropsilus Mik, 1878: 6
Anepsiomyia Bezzi, 1902: 192
Nematoproctus Loew, 1857: 40
Phasmaphleps Bickel, 2005: 21
Terpsimyia Dyte, 1975: 275
Urodolichus Lamb, 1922: 394
Tribe Diaphorini Schiner, 1864
Achradocera Becker, 1922: 207
Aphasmaphleps Grichanov, 2010: 406
Arabshamshevia Naglis, 2014: 728
Asyndetus Loew, 1869: 34
Chrysotus Meigen, 1824: 40
Cryptophleps Lichtwardt, 1898: 491
Diaphorus Meigen, 1824: 32
Dubius Wei, 2012: 611
Emiratomyia Naglis, 2014: 726
Falbouria Dyte, 1980: 223
Lyroneurus Loew, 1857: 38
Melanostolus Kowarz, 1884: 51
Ostenia Hutton, 1901: 34
Shamshevia Grichanov, 2012: 558
Trigonocera Becker, 1902: 57
Tribe Argyrini Negrobov, 1986
Argyra Macquart, 1834: 456
Dactylonotus Parent, 1934: 136
Keirosoma Van Duzee, 1929: 24
Pseudargyra Van Duzee, 1930: 31
Somillus Brethes, 1924: 110
Symbolia Becker, 1921: 210
Subfamily Dolichopodinae Latreille, 1809
Remarks. The genus-group names Arpactodolichopodites, Convexivertex,
Eoeuryopterites and Orbicapitis described by Hong (2002) are unavailable,
because they are based on unavailable species-group names proposed after
1999 without specification of a type depository. Hong everywhere in his
big work (2002) mentioned 2001 as the year of de-scription of his new taxa;
however, I have not found his work with their descriptions pub-lished in
2001.
† Arpactodolichopodites Hong, 2002: 355
† Convexivertex Hong, 2002: 351
† Eoeuryopterites Hong, 2002: 348
† Orbicapitis Hong, 2002: 345
† Leptodolichopodites Hong, 2002: 345
† Prohercostomus Grichanov, 1997: 82
Allohercostomus Yang, Saigusa et Masunaga, 2001: 180
Katangaia Parent, 1933: 12
Pseudohercostomus Stackelberg, 1931: 776
Tribe Dolichopodini Latreille, 1809
Afrohercostomus Grichanov, 2010: 184
Ahercostomus Yang et Saigusa, 2001: 239
Ahypophyllus Zhang et Yang, 2005: 180
Anasyntormon Dyte, 1975: 245
Dolichopus Latreille, 1796: 159
Ethiromyia Brooks et Wheeler, 2005: 491
Gymnopternus Loew, 1857: 10
Hercostomus Loew, 1857: 9
Lichtwardtia Enderlein, 1912: 406
Neohercostomus Grichanov, 2011: 6
Neohercostomus Grichanov, 2011: 6 [subgenus]
Subhercostomus Grichanov, 2011: 22 [subgenus]
Ortochile Latreille, 1809: 289
Parahercostomus Yang, Saigusa et Masunaga, 2001: 176
Poecilobothrus Mik, 1878: 1
Pterostylus Mik, 1878: 4
Setihercostomus Zhang et Yang, 2005: 183
Srilankamyia Naglis, Grootaert et Wei, 2011: 3
Sybistroma Meigen, 1824: 71
Tribe Tachytrechini Negrobov, 1986
Afroparaclius Grichanov, 2006: 29
Afropelastoneurus Grichanov, 2006: 28
Apelastoneurus Grichanov, 2006: 24
Aphalacrosoma Zhang et Yang, 2005: 182
Argyrochlamys Lamb, 1922: 391
Cheiromyia Dyte, 1980: 223
Metaparaclius Becker, 1922: 33
Muscidideicus Becker, 1916: 268
Paraclius Loew, 1864: 97
Pelastoneurus Loew, 1861: 36
Phalacrosoma Becker, 1922: 44
Phoomyia Naglis et Grootaert, 2013
Platyopsis Parent, 1929: 12
Pseudargyrochlamys Grichanov, 2006: 31
Pseudoparaclius Grichanov, 2006: 29
Pseudopelastoneurus Grichanov, 2006: 30
Stenopygium Becker, 1922: 75
Tachytrechus Haliday in Walker, 1851:173
Subfamily Enliniinae Robinson, 1970
Enlinia Aldrich, 1933: 168
Harmstonia Robinson, 1964: 122
Haromyia Runyon, 2015: 590
Subfamily Hydrophorinae Lioy, 1864
Tribe Hydrophorini Lioy, 1864
Abatetia Miller, 1945: 72
Anahydrophorus Becker, 1917: 298
Aphrosylopsis Lamb, 1909: 132
Coracocephalus Mik, 1892: 279
Diostracus Loew, 1861: 43
Diostracus Loew, 1861: 43 [subgenus]
Lagodechia Negrobov et Zurikov, 1996: 632 [subgenus]
Ozmena Ozdikmen, 2010: 265 [subgenus]
Sphyrotarsus Mik, 1874: 342 [subgenus]
Eucoryphus Mik, 1869: 19
Helichochaetus Parent, 1933: 354
Hydatostega Philippi, 1866: 779
Hydrophorus Fallen, 1823: 2
Liancalomima Stackelberg, 1931: 778
Liancalus Loew, 1857: 22
Melanderia Aldrich, 1922: 146
Oedematopiella Naglis, 2011: 47
Oedematopus Van Duzee, 1929: 38
Orthoceratium Schrank, 1803: 55
Paraliancalus Parent, 1938: 213
Paraliptus Bezzi, 1923: 179
Rhynchoschizus Dyte, 1980: 223
Scellus Loew, 1857: 22
Scorpiurus Parent, 1933: 357
Tribe Aphrosylini Aldrich, 1905
Acymatopus Takagi, 1965: 78
Aphrosylus Haliday in Walker, 1851: 220
Cemocarus Meuffels et Grootaert, 1984: 152
Conchopus Takagi, 1965: 49
Cymatopus Kertesz, 1901: 408
Paraphrosylus Becker, 1921: 27
Teneriffa Becker, 1908: 54
Thambemyia Oldroyd, 1956: 210
Thambemyia Oldroyd, 1956: 210 [subgenus]
Prothambemyia Masunaga, Saigusa et Grootaert, 2005 [subgenus]
Thinolestris Grootaert et Meuffels, 1988: 32
Tribe Epithalassiini Grichanov, 2008
Epithalassius Mik, 1891: 186
Peodes Loew, 1857: 29
Tribe Hypocharassini Negrobov, 1981
Hypocharassus Mik, 1879: 627
Tribe Thinophilini Aldrich, 1905
Machaerium Haliday, 1832: 351
Nanothinophilus Grootaert et Meuffels, 1998: 165
Paralleloneurum Becker, 1902: 51
Thinophilus Wahlberg, 1844: 37
Thinophilus Wahlberg, 1844: 37 [subgenus]
Schoenophilus Mik, 1878: 9 [subgenus]
Subfamily Kowmunginae Yang, Zhu, Wang et Zhang, 2006
Kowmungia Bickel, 1987: 147
Phacaspis Meuffels et Grootaert, 1988: 312
Subfamily Medeterinae Lioy, 1864
† Medeterites Grichanov, 2010: 209
† Prosystenus Negrobov, 1976:122
Atlatlia Bickel, 1986: 165
Cryptopygiella Robinson, 1975: 40
Dominicomyia Robinson, 1975: 40
Grootaertia Grichanov, 1999: 90
Hurleyella Runyon et Robinson, 2010
Microchrysotus Robinson, 1964: 251
Microcyrtura Robinson, 1964: 245
Micromedetera Robinson, 1975: 37
Papallacta Bickel, 2006: 211
Pharcoura Bickel, 2007: 6
Pindaia Bickel, 2014: 210
Tribe Medeterini Lioy, 1864
Asioligochaetus Negrobov, 1966: 877
Craterophorus Lamb, 1921: 475 (1922: 380)
Cyrturella Collin, 1952: 142
Demetera Grichanov, 2011: 10
Dolichophorus Lichtwardt, 1902: 199
Medetera Fischer von Waldheim, 1819: 7
Medeterella Grichanov, 2011: 9
Nikitella Grichanov, 2011: 2
Paramedetera Grootaert et Meuffels, 1997: 309
Saccopheronta Becker, 1914: 215
Tribe Systenini Robinson, 1970
† Palaeosystenus Grichanov, Negrobov et Selivanova, 2014: 66
† Systenites Grichanov, Negrobov et Selivanova, 2014: 62
Euxiphocerus Parent, 1935: 121
Systenomorphus Grichanov, 2010: 80
Systenoneurus Grichanov, 2010: 83
Systenus Loew, 1857:34
Tribe Thrypticini Negrobov, 1986
Corindia Bickel, 1986: 137
Thrypticus Gerstaecker, 1864: 43
Tribe Udzungwomyiini Grichanov, 2019
Maipomyia Bickel, 2004: 844
Neomedetera Zhu, Yang et Grootaert, 2007: 227
Udzungwomyia Grichanov, 2018: 10
Subfamily Neurigoninae Aldrich, 1905
Arachnomyia White, 1916: 252
Halteriphorus Parent, 1933: 368
Naticornus Oleinichek, 2005: 247
Oncopygius Mik, 1866: 307
Tribe Neurigonini Aldrich, 1905
Bickelomyia Naglis, 2002: 228
Neurigona Rondani, 1856: 142
Viridigona Naglis, 2003 (2002): 564
Tribe Dactylomiini Naglis, 2002
Argentinia Parent, 1931: 17
Dactylomyia Aldrich, 1894: 151
Macrodactylomyia Naglis, 2002 (2001): 496
Systenoides Naglis, 2002 (2001): 499
Tribe Coeloglutini Naglis, 2001
Coeloglutus Aldrich, 1896: 338
Mberu Capellari et Amorim, 2011: 39
Neotonnoiria Robinson, 1970: 40-17
Paracoeloglutus Naglis, 2001: 200
Subfamily Peloropeodinae Robinson, 1970
Alishanimyia Bickel, 2007: 68
Chrysotimus Loew, 1857: 48
Cremmus Wei, 2006: 490, 500
Discopygiella Robinson, 1965: 51
Fedtshenkomyia Stackelberg, 1927: 230
Griphophanes Grootaert et Meuffels, 1998: 191
Guzeriplia Negrobov, 1968: 470
Hadromerella De Meijere, 1916: 253
Meuffelsia Grichanov, 2008: 160
Micromorphus Mik, 1878: 6
Nanomyina Robinson, 1964: 119
Neochrysotimus Yang, Saigusa et Masunaga, 2008: 20
Nepalomyia Hollis, 1964: 110
Notobothrus Parent, 1931: 19
Peloropeodes Wheeler, 1890: 373
Pseudoxanthochlorus Negrobov, 1977: 146
Vetimicrotes Dyte, 1980: 223
Subfamily Plagioneurinae Aldrich, 1905
Plagioneurus Loew, 1857: 43
Subfamily Rhaphiinae Bigot, 1852
Haplopharyngomyia Meuffels et Grootaert, 1999: 289
Ngirhaphium Evenhuis et Grootaert, 2002: 310
Physopyga Grootaert et Meuffels, 1989: 200
Rhaphium Meigen, 1803: 272
Subfamily Sciapodinae Becker, 1917
† Wheelerenomyia Meunier, 1908: 9
Pouebo Bickel, 2008: 50
Tribe Mesorhagini Bickel, 1994
Amesorhaga Bickel, 1994: 82
Mesorhaga Schiner, 1868: 217
Negrobovia Bickel, 1994: 83
Tribe Sciapodini Becker, 1917
Condylostylus Bigot, 1859: 215
Bickelia Grichanov, 1996: 119
Dytomyia Bickel, 1994: 91
Helixocerus Lamb, 1929: 137
Mascaromyia Bickel, 1994: 100
Narrabeenia Bickel, 1994: 101
Naufraga Bickel, 1992: 36
Pilbara Bickel, 1994: 99
Sciapus Zeller, 1842: 831
Sinosciapus Yang, 2002: 432
Tribe Chrysosomatini Becker, 1918
Abbemyia Bickel, 1994: 212
Amblypsilopus Bigot, 1888: 24
Austrosciapus Bickel, 1994: 231
Bickeliolus Grichanov, 1996: 224
Chrysosoma Guerin-Meneville, 1831: pl.20, fig.6
Chrysosoma Guerin-Meneville, 1831: pl.20, fig.6 [subgenus]
Mesoblepharius Bigot, 1859: 215 [subgenus]
Kalocheta Becker, 1923: 41 [subgenus]
Ethiosciapus Bickel, 1994: 141
Gigantosciapus Grichanov, 1997: 79
Heteropsilopus Bigot, 1859: 215
Krakatauia Enderlein, 1912: 408
Lapita Bickel, 2002: 36
Parentia Hardy, 1935: 245
Plagiozopelma Enderlein, 1912: 367
Pseudoparentia Bickel, 1994: 135
Subfamily Sympycninae Aldrich, 1905
Tribe Sympycnini Aldrich, 1905
Brevimyia Miller, 1945: 72
Calyxochaetus Bigot, 1888: 24
Chaetogonopteron De Meijere, 1914: 96
Colobocerus Parent, 1933: 403
Erebomyia Runyon et Hurley, 2003: 114
Filatopus Miller, 1945: 72
Hercostomoides Meuffels et Grootaert, 1997: 474
Humongochela Evenhuis, 2004: 35
Hyptiocheta Becker, 1922: 245
Ischiochaetus Bickel et Dyte, 1989: 414
Lamprochromus Mik, 1878: 7
Liparomyia White, 1916: 255
Micropygus Bickel et Dyte, 1989: 415
Neoparentia Robinson, 1967: 252
Nothorhaphium Bickel, 1999: 181
Nurteria Dyte et Smith, 1980: 461
Olegonegrobovia Grichanov, 1995: 125
Parasyntormon Wheeler, 1899: 41
Phrudoneura Meuffels et Grootaert, 1987: 319
Pinacocerus Van Duzee, 1930: 11
Scelloides Bickel et Dyte, 1989: 415
Scotiomyia Meuffels et Grootaert, 1997: 248
Suschania Negrobov, 2003: 35
Sympycnus Loew, 1857: 42
Syntormon Loew, 1857: 35
Telmaturgus Mik, 1874: 349
Tetrachaetus Bickel et Dyte, 1989: 417
Teuchophorus Loew, 1857: 44
Yumbera Bickel, 1992: 1007
Tribe Stolidosomatini Becker, 1922
Pseudosympycnus Robinson, 1967: 901
Stolidosoma Becker, 1922: 213
Sympycnidelphus Robinson, 1964: 149
Tribe Campsicnemini Becker, 1917 + Eurynogaster complex
Adachia Evenhuis, 2005: 44
Arciellia Evenhuis, 2005: 45
Campsicnemus Haliday in Walker, 1851: 187
Elmoia Evenhuis, 2005: 48
Eurynogaster Van Duzee, 1933: 339
Major Evenhuis, 2005: 51
Sigmatineurum Parent, 1938: 215
Sweziella Van Duzee, 1933: 346
Uropachys Parent, 1935: 80
Subfamily Tenuopodinae Grichanov, 2018
Tenuopus Curran, 1924: 228
Subfamily Xanthochlorinae Aldrich, 1905
Xanthochlorus Loew, 1857: 42