Afrotropical species of the genus Condylostylus Bigot
(Diptera: Dolichopodidae)
Igor Ya. GRICHANOV
Grichanov, I.Ya. Afrotropical species of the genus
Condylostylus Bigot (Diptera: Dolichopodidae)
C. galinae sp.n. from Uganda, male of C. paricoxa Parent
from Kenya are described. Catalogue and a key to 12 known
species of Condylostylus are given.
I.Ya. Grichanov, All-Russian Institute of Plant Protection,
Podbelskogo 3, St.Petersburg-Pushkin, 189620, Russia.
Key words: Diptera, Dolichopodidae, Condylostylus,
Tropical Africa.
Introduction
The genus Condylostylus is a wide spread pantropical taxon. It
contains approximately 308 described species with a huge diversity
in Neotropical Region (230 species) and a small number of species in
southern Nearctic, far-eastern Palearctic and other Regions (Bickel,
1994). Twelve species of the genus occur throughout Africa south of
latitude 10 centigrades North including Madagascar. Probably a few
afrotropical species of Condylostylus await description.
The species of Condylostylus, contrary to that of other
sciapodine genera, are well recognized by strongly recurved vein M1,
four strong scutellar and five pairs of strong dorsocentral bristles,
and setose mound on the frons, bearing vertical seta. Males of
afrotropical species of the genus sometimes have abnormal wing
venation (C. pateraeformis Group), very narrow face and greatly
reduced hairs on the frontal mound (the same Group and C. burgeoni
Group), relatively short (C. paricoxa) or bifurcated (C. selectus)
cercus.
In this paper first descriptions of C. galinae sp.n. and male C.
paricoxa Parent, and new records for known species are given on the
basis of the collections from the Natural History Museum, London
(NHML), the Hungarian Natural History Museum (HNHM), and the Institut
Royal des Sciences Naturelles, Brussels (IRSN).
Holotype and paratypes of the new species, as well as a male of
C. paricoxa, are conserved in the Natural History Museum (London).
List of known species of Condylostylus
(for references see Dyte & Smith, 1980)
Condylostylus Bigot, 1859:215. Type species Psilopus bituberculatus
Macquart, 1842, original designation.
angustipennnis Loew, 1858:372 (1860:346) (Psilopus) - South Africa.
beckeri Speiser, 1920:218 - Cameroun.
burgeoni Parent, 1935:115 - Zaire, Tanzania, Kenya (!), Ruanda (!).
congensis Curran, 1927:263 - Congo, Cameroun, Zaire, South
Africa, (?) Madagascar, Uganda (!), Tanzania (!).
degener Parent, 1934:119 - South Africa.
galinae sp.n. - Uganda.
imitator Curran, 1924:221 - Zaire, Kenya, Tanzania, Malawi,
Zimbabwe, Mozambique, South Africa, Angola (!), Namybia (!).
= imitans Curran, 1925:114 (replacement).
kivuensis Vanschuytbroeck, 1964:136 - Zaire.
paricoxa Parent, 1939:267 - Kenya, Zimbabwe, Tanzania (!).
pateraeformis Becker, 1923:38 - Cameroun, Nigeria, Zaire, Uganda,
Tanzania, South Africa, (?) Madagascar, Kenya (!).
= alter Becker, 1923:38.
selectus Parent, 1931:43 - Malawi, Zaire.
stenurus Loew, 1858:377 (1860:346) (Psilopus) - South Africa,
Zimbabwe (!).
= sicatris Curran, 1926:389.
Key to known species of Condylostylus
Males
1. Fore tibia with two long apicoventral setae, half as long as fore
basitarsomere.......................................C. paricoxa Parent
- Fore tibia without such setae, at most with a long apicoventral
hair.................................................................2
2. Cercus bifurcated, with long narroe branches.....C. selectus Parent
- Cercus not bifurcated, usually long, sometimes short, with
prominence...........................................................3
3. Middle tibia with a row of long dorsal setae, femora bare.........4
- Middle tibia with a few usual anterodorsal and posterodorsal
bristles, or femora with long hairs..................................5
4. Cercus nearly twice as long as 7th tergum and epandrium together...
....................................................C. burgeoni Parent
- Cercus nearly thrice as long as 7th tergum and epandrium together..
......................................................C. galinae sp.n.
5. Hind tibia with black ventral cicatrix (callus-like swelling) in
basal third, covered with light hairs.................C. stenurus Loew
- Hind tibia without such cicatrix..................................6
6. Pedicel with long ventral and dorsal setae, as long as three
antennal segments together; costa with curvation in front of R1;
posterior wing edge sinuous......................C. angustipennis Loew
- Pedicel with usual setae, at most as long as scape and pedicel
together; costa without curvation....................................7
7. Wing with normal female-type venation.............C. degener Parent
- Venation abnormal: M1+2 curved towards posterior wing border, M1
continued nearly in the same line as M2..............................8
8. Fore tibia enlarged, fore tarsus simple..........C. beckeri Speiser
- Fore tibia simple, fore basitarsomere enlarged....................9
9. M1+2 and M1 form acute angle................C. pateraeformis Becker
- M1+2 and M1 form right angle.....................................10
10. Fore coxa brown, middle and hind coxae black, legs brown, first
tarsomere of fore tarsus longer and wider than in C. congensis and C.
imitans, cercus longer, than in those species.........................
...........................................C. kivuensis Vanshuytbroeck
- Other combination of features...................................11
11. Fore coxa black at least in basal half; fore femora brown-black...
...................................................C. congensis Curran
- Fore coxa and femora yellow.....................C. imitator Curran
Females
1. Middle coxa mostly yellow........................C. paricoxa Parent
- At least middle coxa mostly brown or black........................2
2. Squamae with black ciliae..........................C. stenurus Loew
- Squamae with yellow ciliae........................................3
3. Wing anal lobe and angle absent....................................
..................................C. burgeoni Parent, C. galinae sp.n.
- Wing anal lobe present, anal angle obtuse or acute................4
4. Hind femora yellow, at most with black spot on apex................
....................................................C. imitator Curran
- Hind femora partly black..........................................5
5. Frons blue-violet...........................C. pateraeformis Becker
- Frons green.....................................C.
congensis Curran
Descriptions and new records
Condylostylus galinae sp.n. (Fig. 1)
Holotype. Male. Uganda: Ruwenzori Range, XII.1934-I.1935. B.M.E.Afr.
Exp. B.M.1935-203 / Namwamba Valley, 6500 ft (F.W.Edwards). Paratypes.
2 males and a female, the same labels.
Description. Frons metallic blue-green, mostly white pollinose.
A strong front vertical bristle bends forward, arising from small
mound; this mound usually bare, at most with one fine dark hair;
postvertical bristle is positioned as a linear continuation of the
postocular setal row. Ventral postcranium covered with irregular
white hairs. Face black, silvery-white pollinose, narrow, 10 times as
high as wide in the middle. Bulging clypeus nearly half as wide as
epistome under antennae. Proboscis orange-brown, palpi black, with
numerous light hairs. Antennae black, as long as height of head.
Pedicel with short dorsal and ventral bristles. First flagellomere
rounded, as long as its height, densely pubescent. Arista dorsal,
microscopically haired. Length ratio of scape to pedicel to first
flagellomere to arista - 7:6:8:85.
Mesonotum and scutellum brilliantly shining green-violet. Pleura
bronze-green, white pollinose. Dorsocentral bristles gradually
decreasing in size anteriorly with two strongest posterior pair.
Short acrostichals in two rows, restricted to anterior half of
mesonotum. Scutellum with two pair of strong bristles, with lateral
setae somewhat smaller.
Legs dirty-yellow. Middle and hind coxae dark-brown, whitish
pollinose, middle and hind femora more or less brownish in apical
half, hind tibia brown, hind tarsus and apical segments of other
tarsi black. Fore coxa from the front with numerous yellow hairs.
Middle and hind coxae from the outside with a few yellow hairs.
Femorae without strong or long bristles and hairs. Middle tibia with
a posterodorsal bristle. Fore tibia with a long black apicoventral
hair. Fore basitarsomere flattened in apical half. Length ratio of
fore coxa to femora to tibia to tarsus (segments from first to fifth)
- 1.7 : 2.6 : 3.2 : 2.4 : 0.8 : 0.6 : 0.4 : 0.2. Middle tibia
slightly flattened in basal two thirds, with 8-9 long black dorsal
hairs in middle half, with irregular ventral pectination in second
quarter. Fourth tarsomere enlarged, with a few long black dorsal
setae. Fifth tarsomere with a brush of short erected ventral hairs.
Length ratio of middle coxa to femora to tibia to tarsus (segments
from first to fifth) - 1.4 : 4.0 : 6.0 : 4.3 : 0.7 : 0.4 : 0.3 : 0.9.
Last tarsomeres of hind tarsi slightly thickend. Length ratio of hind
coxa to femora to tibia to tarsus (segments from first to fifth) -
1.0 : 5.3 : 7.1 : 3.7 : 1.0 : 0.7 : 0.5 : 0.4.
Wings hyaline, slightly darkened in anterior part, veins brown.
R4+5 gently curved to M1 in apical fifth. M1 slightly curved basad.
Ratio of parts of costa between R2+3 and R4+5 to those between R4+5
and M1 - 45 : 7. Crossvein m-cu straight. Ratio of crossvein m-cu to
apical part of M1+2 (fork-handle) to apical part of CuA - 43 : 63 :
18. Anal vein and lobe reduced. Anal angle absent. Squamae yellow,
with brown apex and pale bristles. Halteres yellow with brown knob,
halter stem thin and long.
Abdomen thin and long, mostly black, with short black hairs.
First two segments and apical border of third to sixth segments
metallic green-violet; first tergum with broad membranous excavation
and long white lateral hairs. Unmodified segments together nearly 4
times as long as mesonotum. Hypopygium black, with short black hairs.
Cercus long, filiform, brown, laterally with a row of long black
bristles in basal half, ventrally densely setose with short curved
light hairs. Cercus nearly thrice as long as seventh tergum and
epandrium together. Surstylus and epandrial lobe greatly reduced.
Female. Similar to male except lacking male secondary sexual
characters, otherwise as follows: face 5 times as high as wide in the
middle. Legs mostly yellow, middle and hind coxae, hind tibia and knee
brownish, hind tarsus and apical tarsomeres of other tarsi dark-brown.
All coxae with additional 1-2 dark hairs. Middle tibia with one
anterodorsal and two posterodorsal bristles. Ratio of first to second
tarsomere of fore, middle and hind legs - 9.5:2.5; 12.5:3.0; 10.3:3.5.
Wing with indistinct dark spot of shape usual for genus. Ratio of m-cu
to apical part of M1+2 - 43:81. Halters yellow. Abdomen comparatively
short, entirely shining blue-green.
Length: male body 8.4 mm; antenna 1.5 mm; postabdomen 2.6 mm;
female body 6.1 mm; wing-length 5.6 mm; wing-width 1.7 mm.
Distribution: West Uganda.
Etymology. The species is named for Ms Galina Grichanova, who had
the scrupulous and subtle job of illustrating many authors papers.
Diagnosis. Males of C. galinae are closely related to C. burgeoni
and can be separated by the following combination of attributes: fore
femora, fore and middle tibiae yellow, middle and hind femorae mostly
yellow; wing fork-handle 1.5 times as long as m-cu; cercus nearly
thrice as long as seventh tergum and epandrium together, with long
bristles in basal half. Female probably similar to C. burgeoni.
Condylostylus burgeoni Parent (Fig. 2)
Material examined. 2 males & 4 females, Kenya: 18-20.XII.1970,
A.E. Stubbs, B.M. 1972-211 / Kakamega Forest, 5200 feet; 2 males,
Cent. Africa: Ruanda Urundi, X.1956, N.L.H. Krauss. B.M. 1957-4.
Diagnosis. C. burgeoni is closely related to C. galinae sp.n.,
differing by brown-black middle and hind legs and quantitative
characters such as follows: apical part of M1+2 (fork-handle) at
least twice as long as m-cu; cercus nearly twice as long as 7th
tergum and epandrium together, with long setae from base up to the
apex. Frons usually bare in both sexes, with strong vertical seta and
at most with one fine hair on small mound. Despite the description by
Parent, wing darkened in anterior part, with indistinct spot of usual
for genus shape. Females differ from males by yellow middle femora,
middle tibia and (except knee) hind femora, also by darker spot on
wing.
Distribution: Zaire, Tanzania, Kenya (!), Ruanda (!).
Condylostylus paricoxa Parent (Fig. 3)
Material examined. 1 male & 3 females, Kenya: Diani Beach,
VII.1951, N.L.H.Krauss. B.M. 1951-541; 2 females, Kenya: Teita Hills,
VI.[19]48, van Someren / Com. Inst. Ent. Coll. No. 13261; 1 female,
Tanganyika: Amani, 1957, J.G. Halcrow, in forest (20) / C.I.E. Coll.
No. 15425 / Pres. by Com. Inst. Ent. B.M. 1969-3.
Description (male). Corresponds to female described by Parent
(1939) with the exception of secondary sexual characters and other
features as noted. Frons metallic blue-violet, slightly pollinose,
with strong front vertical bristle and white hairs, arising from
distinct mound. Face blue-black, silvery-white pollinose, 3 times as
high as wide in the middle. Bulging clypeus nearly half as wide as
epistome under antennae. Scape and pedicel short, dark yellow, first
flagellomere absent.
Mesonotum and scutellum brilliantly blue-green. Pleura brown
with metallic reflection, white pollinose. 5 long dorsocentral
bristles with a hair in front of the first one, 3 pairs of
acrostichals.
Legs pale-yellow, hind tarsus and apical segments of other tarsi
brown. Fore coxa from the front with numerous yellow hairs and 3
preapical black bristles. Middle coxa from the outside with a few
yellow and dark hairs, hind coxa with a black seta. Fore and middle
femorae with fine yellow ventral hairs, as long as femora diameter.
Fore tibia with a long black preapical ventral hair, with 2 long
thick black apicoventral setae, half as long as fore basitarsomere.
First tarsomere slightly swollen in middle half, with short dense
ventral hairs and one thin apicodorsal seta. Length ratio of fore
coxa to femora to tibia to tarsus (segments from first to fifth) - 51
: 85 : 97 : 68 : 33 : 17 : 12 : 10. Middle tibia with 3 strong
anterodorsal and 3 weak posterodorsal bristles. Length ratio of
middle coxa to femora to tibia to tarsus (segments first and second)
- 35 : 115 : 160 : 114 : 27. Last tarsomeres of hind tarsi slightly
flattened. Length ratio of hind coxa to femora to tibia to tarsus
(segments from first to fifth) - 35 : 125 : 138 : 112 : 30 : 20 : 13
: 9.
Wing with a spot as figured by Parent (1939). R4+5 gently curved
to M1 in apical fifth. M1 slightly curved basad. Ratio of parts of
costa between R2+3 and R4+5 to those between R4+5 and M1 - 47 : 4.
Crossvein m-cu straight. Ratio of crossvein m-cu to apical part of M1
+2 (fork-handle) to apical part of CuA (up to the wing border) - 33 :
79 : 20. Anal vein reduced. Anal lobe present. Squamae yellow, with
brown apex and brown-black bristles. Halteres yellow, halter stem
thin and long.
Abdomen comparatively short, metallic green-violet, with black
hairs. First tergum with narrow membranous excavation and short white
lateral hairs. Unmodified segments together nearly 1.5 times as long
as mesonotum. Hypopygium small, brown. Cercus yellow, short,
spoon-shaped, curved, especially on apex, with yellow hairs.
Surstylus very short, narrow, epandrial lobe greatly reduced, with
strong setae.
Length: male body 4.2 mm; wing-length 4.7 mm; wing-width 1.5 mm.
Distribution: Kenya, Zimbabwe, Tanzania (!).
Diagnosis. C. paricoxa is the only species in Africa with two
long preapical setae on fore tibia. It is close to C. victoriseta
(Hollis, 1964) from Sumatra and can be separated by the following
combination of attributes: frons with white hairs; fore tibia with a
long black preapical ventral hair, with 2 long thick black
apicoventral setae of equal length, half as long as fore
basitarsomere. First tarsomere slightly swollen in middle half, with
short dense ventral hairs and one thin apicodorsal seta. Cercus
yellow, short, spoon-shaped, curved, especially on apex, with yellow
hairs. Surstylus greatly reduced, narrow, epandrial lobe very short,
with strong setae.
Condylostylus selectus Parent
Material examined. 1 male & 3 females, Zaire: Park National
Albert, 1953-1954 (P. Vanschuytbroeck, J. Kekenbosch, H. Synave)
[IRSN].
Diagnosis. Males of C. selectus can be easily identified by
bifurcated cercus with long narrow branches.
Distribution: Malawi, Zaire.
Condylostylus stenurus (Loew)
Material examined. 1 male & 1 female, S. Rhodesia: Kessesse R.
[&] Nyanyana R. area, Tsetse Fly Ops., 18 [&] 14.1.1956, 4 [&] 10 m.
E of Kanba Gorge, Coll. R. Goovier (158 [&] 146) [NHML].
Diagnosis. C. stenurus is the only afrotropical species with
ventral callus-like swelling on hind tibia. It has small frontal mound
with strong vertical seta and a few microscopic hairs. Legs and coxae
mostly yellow, middle coxa anteriorly mostly brown; wing with
female-type venation; cercus 1.5 times as long as 7th tergum and
epandrium together, with small basoventral projection.
Distribution: South Africa, Zimbabwe
(!).
Condylostylus pateraeformis Becker
Material examined. 5 males & 9 females, Uganda: Ruwenzori Range,
XII.1934-I.1935. B.M.E.Afr. Exp. B.M. 1935-203 / Kilembe, 4500 ft (F.
W.Edwards); 1 male, Uganda: Entebbe, 13.XII.1934, F.W. Edwards. B.M.
1935-203; 1 male, Uganda: Lake near R. Muzizzi, Day 2.XI.1912, Dr. R.
E. McConnell; 1 female, Uganda: Ruwenzori Range, Bundibugyo, 3440
ft., 22.VIII-3.XI.1952, D.S. Fletcher / Ruwenzori Exped. B.M.
1952-566. 1 female, Uganda: Acholi Dist., Gulu, 19.X.1964, R.W.
Crosskey; 10 males, van Someren, Bwamba, 4-44 / V.G.L. van Someren
Collection, Brit. Mus. 1959-468; 1 male & 4 females, van Someren,
Jombeni Hills, Kenya, 5-47 / Com. Inst. Ent. Coll. No. 13261; 3
females, van Someren, Nyeri (S.), Kenya, 1948 / Com. Inst. Ent. Coll.
No. 11464; 1 female, Kawanga, on cassava leaf, 11.V.1950 / Coll.No.
1745, Ident. No. / Com. Inst. Ent. Coll. No. 14896; 3 females, Kenya:
Kwali Forest, 20 mls W of Mombasa, 1.VI.1948 / on edge of forest /
Miss. M. Steele. B.M. 1948-347; 4 females, Kenya: 15-16.XII.1970,
A.E. Stubbs. B.M. 1972-211 / Lake Nakuru, 5767 feet; 2 males,
Tanganyika: Amani, 1957, J.G. Halcrow, in cattle shed (51) / C.I.E.
Coll. No. 15455 / Pres. by Com. Inst. Ent. B.M. 1969-3; 1 female,
Tanzania: 19[67?]. Ex. Coll. W.H. Potts. B.M. 1967-472; 1 male, Gold
Coast: Aburi, 1912- 13, W.H. Patterson; 1 male: N.Nigeria: Kagoro
Forest, 7.V.1975, J.C. Deeming; 1 female, N.Nigeria: Niger Prov.,
Mokwa, Zugurma, 29.VII.1971, J.C. Deeming, kurmi [NHML].
Additional material. 372 males and females, Congo Belg.: Park
National Albert, 1953-1958 (P. Vanschuytbroeck, V. Hendricks, J.
Kekenbosch, H. Synave) [IRSN]; 3 males, Congo Belg.: Rutshuru, 1937
(J. Ghesquiere) [IRSN]; 9 males and females, Congo Belg.: Bambesa,
1938 (J. Vrijdagh) [IRSN].
Diagnosis. Frons with strong vertical bristle arising from small
bare mound; face 8 times as high as wide in the middle; legs mostly
brown-black, fore coxa, femora and tibia and usually middle tibia
(except apex) yellow; ratio of fore tibia to length of basitarsomere
to width of the same joint to second tarsomere - 115 : 55 : 15 : 35;
M1+2 and M1 form the acute angle; cercus thin, nearly thrice as long
as 7th tergum and epandrium together, with oval ventral lobe.
Distribution: Cameroun, Nigeria, Zaire, Uganda, Tanzania, South
Africa, (?) Madagascar, Kenya (!).
Condylostylus imitator Curran
Material examined. 1 female, Kenya: Diani Beach, VIII.1951, N.L.
H.Krauss. B.M. 1951-541; 1 female, Tanganyika: Amani, 1954, J.G.
Halcrow, in forest (20) / C.I.E. Coll. No. 15425 / Pres. by Com.
Inst. Ent. B.M. 1969-3; 1 male, Tanzania: Mituni Nat. Park /
4.XI.1987, leg. Mahunka [HNHM]; 1 male, S.W. Africa (W50): Gobiswater
Fm., 12 mls. N, Grootfontein, 5.VI.1972 / Southern African Exp. B.M.
1972-1; 1 female, Portug. E.Afr.: E. of Mt. Mlanje, 28.XI.1913, S.A.
Neave / 28.XI.13, S.A.N.; 1 female, S.Africa: Cape Province, Durban,
21.XI.1954, C.H. Andrews / Brit. Mus. 1955-68; 1 female, Mozambique:
Machava, 5 m.N. Lourenco, Marques, 28.XI.1955 / I.R. James & L.F.
Brown. B.M. 1955-471; 4 females, Angola (A11): Bruco,
26.XII-2.III.1972 / Southern African Exp. B.M. 1972-1.
Additional material. 1 male, Congo Belg.: Park National Albert,
1955 (P. Vanschuytbroeck) [IRSN]; 3 males, Congo Belg.: Rutshuru,
1937 (J. Ghesquiere) [IRSN].
Diagnosis. Similar to C. pateraeformis except as noted. Face 7
times as high as wide in the middle; legs mostly yellow, middle and
hind coxae, 2nd-5th tarsomeres of fore tarsus, apex of middle tibia,
middle and hind tarsi, middle femora in apical fifth black-brown,
hind tibia brown; ratio of fore tibia to length of basitarsomere to
width of the same joint to second tarsomere - 95 : 50 : 12 : 28; M1+2
and M1 form the right angle; M1 with distinct elbow; cercus thin,
nearly thrice as long as 7th tergum and epandrium together, ventral
lobe of cercus with pointed apex.
Distribution: Zaire, Kenya, Tanzania, Malawi, Zimbabwe,
Mozambique, South Africa, Angola (!), Namybia (!).
Condylostylus congensis Curran
Material examined. 2 males, Uganda: Ruwenzori Range, XII.1934-I.
1935. B.M.E.Afr. Exp. B.M. 1935-203 / Kilembe, 4500 ft (F.W.Edwards);
2 males, Zanzibar: Mangapwani, XI.1951, N.L.H. Krauss.
Diagnosis. Similar to C. pateraeformis except as noted. Face 9
times as high as wide in the middle; legs mostly brown-black, fore
coxa in apical half and fore tibia yellow; fore femora and
basitarsomere and sometimes middle tibia yellowish-brown; ratio of
fore tibia to length of basitarsomere to width of the same joint to
second tarsomere - 98 : 53 : 13 : 25; M1+2 and M1 form the right
angle; M1 nearly straight; cercus nearly twice as long as 7th tergum
and epandrium together, with egg-shaped ventral lobe.
Distribution: Congo, Cameroun, Zaire, South Africa, (?)
Madagascar, Uganda (!), Tanzania (!).
Acknowledgements
I am sincerely grateful to Dr. Brian Pitkin, Dr. Laslo Papp, Dr.
P.Vanschuytbroeck and Dr. Oleg Negrobov for their kindness in giving
me the opportunity to study the collections of the Natural History
Museum (London), the Hungarian Natural History Museum (Budapest) and
the Institut Royal des Sciences Naturelles (Brussel). I appreciate
the help I have received from Ms Galina Grichanova in doing the
illustrations.
References
Bickel D.J. 1994. The Australian Sciapodinae (Diptera:
Dolichopodidae), with a review of the Oriental and Australasian
faunas, and a world conspectus of the subfamily. - Rec. Austral.
Mus., 1994, Suppl. 21: 1-394.
Dyte C.E. & Smith K.G.V. 1980. Family Dolichopodidae. In R.W.
Crosskey (ed.). Catalogue of the Diptera of the Afrotropical Region.
Brit. Mus. (Nat. Hist.), London, p. 443-463.
Hollis D. 1964. Notes and descriptions of Indonesian
Dolichopodidae in the Zoologisch Museum, Amsterdam. - Beaufortia, 10,
129, p. 239-274.
Parent O. 1939. Dipteres Dolichopodides de la region ethiopienne.
- Rev. Zool. Bot. Afr., 1939, 32, 2, p. 256-282.
Remark under figures
Figs 1-3. Hypopygium, lateral veiw. 1, C. galinae sp.n.; 2, C.
burgeoni Parent; 3, C. paricoxa Parent.